Exploring evolutionary change of complex phenotypes across time, space, and contexts 
using an integrative approach.
themes, approaches, and systems
Bio-invasions as 'natural' experiments to study (human-induced) contemporary evolution
Islands are nature's test tubes  — 
​simple systems with multiple replicates.
Phylogenetic comparative methods to explore the history of organismal evolution and diversification. 

​Bio-imaging techniques to capture morphological variation in detail
Implementing evolutionary principles into conservation management